/* ============================================================
   IONIS Research Papers — Frontiers-style article layout
   Scoped to .paper-article wrapper (papers section only)
   ============================================================ */

/* Article container — constrain width, center */
.paper-article {
  max-width: 820px;
  margin: 0 auto;
  line-height: 1.75;
}

/* --- Header Block --- */
.paper-header {
  border-bottom: 3px solid var(--md-primary-fg-color);
  padding-bottom: 1.5rem;
  margin-bottom: 2rem;
}

.paper-header .paper-title {
  font-size: 2rem;
  font-weight: 700;
  line-height: 1.2;
  margin: 0 0 0.25rem 0;
  color: var(--md-default-fg-color);
}

.paper-header .paper-subtitle {
  font-size: 1.15rem;
  font-weight: 400;
  font-style: italic;
  color: var(--md-default-fg-color--light);
  margin: 0 0 1.25rem 0;
}

.paper-header .paper-authors {
  font-size: 0.95rem;
  margin: 0 0 0.25rem 0;
  color: var(--md-default-fg-color);
}

.paper-header .paper-affiliation {
  font-size: 0.85rem;
  color: var(--md-default-fg-color--light);
  margin: 0 0 0.5rem 0;
}

.paper-header .paper-meta {
  display: flex;
  flex-wrap: wrap;
  gap: 1rem;
  font-size: 0.8rem;
  color: var(--md-default-fg-color--lighter);
  margin-top: 0.75rem;
}

.paper-header .paper-meta span {
  display: inline-flex;
  align-items: center;
  gap: 0.3rem;
}

/* --- Abstract Block --- */
.paper-abstract {
  background: var(--md-code-bg-color);
  border-left: 4px solid var(--md-primary-fg-color);
  padding: 1.25rem 1.5rem;
  margin: 0 0 2rem 0;
  border-radius: 0 4px 4px 0;
}

.paper-abstract .abstract-label {
  font-size: 0.85rem;
  font-weight: 700;
  text-transform: uppercase;
  letter-spacing: 0.05em;
  color: var(--md-primary-fg-color);
  margin: 0 0 0.5rem 0;
}

.paper-abstract p {
  font-size: 0.92rem;
  line-height: 1.7;
  margin: 0;
  color: var(--md-default-fg-color);
}

/* --- Keywords --- */
.paper-keywords {
  display: flex;
  flex-wrap: wrap;
  gap: 0.4rem;
  margin: 0 0 2.5rem 0;
}

.paper-keywords .kw-label {
  font-size: 0.8rem;
  font-weight: 700;
  color: var(--md-default-fg-color--light);
  margin-right: 0.25rem;
  align-self: center;
}

.paper-keywords .kw-tag {
  font-size: 0.75rem;
  background: var(--md-code-bg-color);
  border: 1px solid var(--md-default-fg-color--lightest);
  border-radius: 3px;
  padding: 0.15rem 0.5rem;
  color: var(--md-default-fg-color--light);
}

/* --- Section headings — article style --- */
.paper-article h2 {
  font-size: 1.4rem;
  font-weight: 700;
  margin-top: 2.5rem;
  padding-top: 1rem;
  border-top: 1px solid var(--md-default-fg-color--lightest);
}

.paper-article h3 {
  font-size: 1.15rem;
  font-weight: 600;
  margin-top: 1.75rem;
}

.paper-article h4 {
  font-size: 1rem;
  font-weight: 600;
  margin-top: 1.25rem;
}

/* --- Body text refinements --- */
.paper-article p {
  text-align: justify;
  hyphens: auto;
}

.paper-article blockquote {
  border-left: 3px solid var(--md-primary-fg-color);
  font-style: italic;
  color: var(--md-default-fg-color--light);
}

/* --- Citation block at bottom --- */
.paper-citation {
  background: var(--md-code-bg-color);
  border: 1px solid var(--md-default-fg-color--lightest);
  border-radius: 4px;
  padding: 1rem 1.25rem;
  margin-top: 3rem;
  font-size: 0.85rem;
}

.paper-citation .cite-label {
  font-weight: 700;
  font-size: 0.8rem;
  text-transform: uppercase;
  letter-spacing: 0.05em;
  color: var(--md-primary-fg-color);
  margin: 0 0 0.5rem 0;
}

.paper-citation p {
  margin: 0;
  text-align: left;
  line-height: 1.6;
}

/* --- Separator between header area and body --- */
.paper-body-start {
  border: none;
  border-top: 1px solid var(--md-default-fg-color--lightest);
  margin: 0 0 2rem 0;
}